Cagra.1372s0011 (Capsella grandiflora)
bZIP

TF Information

Pfam ID Interpro ID Gene ID CIS-BP ID Sequence source
PF00170 (bZIP_1) IPR011616 Cagra.1372s0011 T071486_2.00 JGI:Phytozome (2018-Apr-12)

Directly determined binding motifs

Name/Motif ID Species Forward Reverse Type/Study/Study ID SR
Score
DBD
Identity
No direct experiments

Motifs from related TFs

Name/Motif ID Species Forward Reverse Type/Study/Study ID SR
Score
DBD
Identity
GBF2
M01781_2.00
Arabidopsis thaliana
BMCASSNNN

NNNSSTGKV
PBM
Weirauch et al.(2014)
pTH7258
0.866 0.922
Q40645_ORYSA
M10050_2.00
Oryza sativa Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$OSBZ8_Q6
0.855 0.703
Q41109_PHAVU
M10078_2.00
Phaseolus vulgaris Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$ROM_Q2
0.853 0.672
PK09653.1
M01156_2.00
Cannabis sativa
NNMCASVNNN

NNNBSTGKNN
PBM
Lambert et al.(2019)
pTH10114
0.850 0.656
TAF1_TOBAC
M10079_2.00
Nicotiana tabacum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$TAF1_01
0.837 0.703
TAF1_TOBAC
M10080_2.00
Nicotiana tabacum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$TAF1_Q2
0.837 0.703
GBF3
M01779_2.00
Arabidopsis thaliana
NBMCASSTVD

HBASSTGKVN
PBM
Weirauch et al.(2014)
pTH7328
0.832 0.734
GBF3
M06770_2.00
Arabidopsis thaliana
DNHDNHSACGTGKMD

HKMCACGTSDNHDNH
Dap-seq
OMalley et al.(2016)
GBF3_colamp
0.832 0.734
GBF3
M06771_2.00
Arabidopsis thaliana
DDWKVYSACGTGGCA

TGCCACGTSRBMWHH
Dap-seq
OMalley et al.(2016)
GBF3_col
0.832 0.734
GBF1
M01783_2.00
Arabidopsis thaliana
HBACGTGKMN

NKMCACGTVD
PBM
Weirauch et al.(2014)
pTH7259
0.808 0.594
GBF1
M09913_2.00
Arabidopsis thaliana Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$GBF1_01
0.808 0.594
GBF1
M09914_2.00
Arabidopsis thaliana Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$GBF1F_Q2
0.808 0.594
GBF1
M09915_2.00
Arabidopsis thaliana Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$GBF1_Q2_01
0.808 0.594
GBF1
M09916_2.00
Arabidopsis thaliana Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$GBF1_Q2
0.808 0.594
BZIP16
M06764_2.00
Arabidopsis thaliana
DDWKVYSACGTGGCA

TGCCACGTSRBMWHH
Dap-seq
OMalley et al.(2016)
bZIP16_colamp_a
0.787 0.531
BZIP16
M06765_2.00
Arabidopsis thaliana
DDWKVHSACGTGGCA

TGCCACGTSDBMWHH
Dap-seq
OMalley et al.(2016)
bZIP16_col_v3a
0.787 0.531
BZIP68
M01770_2.00
Arabidopsis thaliana
BMCASGBND

HNVCSTGKV
PBM
Weirauch et al.(2014)
pTH8192
0.786 0.547
BZIP68
M06752_2.00
Arabidopsis thaliana
DNWKNHSACGTGGCR

YGCCACGTSDNMWNH
Dap-seq
OMalley et al.(2016)
bZIP68_col_a
0.786 0.547
PK23964.1
M01160_2.00
Cannabis sativa
DNKMCACGTV

BACGTGKMNH
PBM
Lambert et al.(2019)
pTH10151
0.786 0.609
CPRF3_PETCR
M10071_2.00
Petroselinum crispum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$CPRF3_01
0.785 0.484
CPRF3_PETCR
M10072_2.00
Petroselinum crispum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$CPRF3_Q2
0.785 0.484
CPRF1_PETCR
M10066_2.00
Petroselinum crispum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$CPRF1_01
0.784 0.609
CPRF1_PETCR
M10067_2.00
Petroselinum crispum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$CPRF1_Q2
0.784 0.609
CPRF1_PETCR
M10068_2.00
Petroselinum crispum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$CPRF_Q2
0.784 0.609
EmBP-1
M02659_2.00
Triticum aestivum
CCACGTSW

WSACGTGG
SELEX
Mathelier et al.(2014)
MA0128.1
0.783 0.609
EMBP1_WHEAT
M09921_2.00
Triticum aestivum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$EMBP1_02
0.783 0.609
EMBP1_WHEAT
M09922_2.00
Triticum aestivum Transfac license required
Transfac license required
Transfac
Matys et al.(2006)
P$EMBP1_Q2
0.783 0.609
For this family, TFs with SR scores > 0.782 will likely have a similar motif

DNA Binding Domains

Protein ID Domain From To Sequence
Cagra.1372s0011.1.p bZIP 247 310

Links

Other bZIP family TFs
Other Capsella grandiflora TFs

228 Related TFs

Name Species Gene ID Motif Evidence SR
Score
Action
Bostr.7365s0064 Boechera stricta Bostr.7365s0064 I 0.886
Carubv10001247m.g Capsella rubella Carubv10001247m.g I 0.886
Thhalv10028761m.g Eutrema salsugineum Thhalv10028761m.g I 0.885
Bo9g003700 Brassica oleracea Bo9g003700 I 0.885
Bra037382 Brassica rapa Bra037382 I 0.885
GBF2 Arabidopsis lyrata Al_scaffold_0006_3899 I 0.882
Gorai.008G123200 Gossypium raimondii Gorai.008G123200 I 0.869
TCM_005210 Theobroma cacao TCM_005210 I 0.869
Gorai.004G153100 Gossypium raimondii Gorai.004G153100 I 0.869
GBF2 Arabidopsis thaliana AT4G01120 D 0.866
GLYMA03G41590 Glycine max GLYMA03G41590 I 0.859
GLYMA19G44190 Glycine max GLYMA19G44190 I 0.859
Sb03g030050 Sorghum bicolor Sb03g030050 I 0.857
OLONG_003667 Oryza longistaminata OLONG_003667 I 0.855
OMERI01G23000 Oryza meridionalis OMERI01G23000 I 0.855
OGLUM05G27920 Oryza glumaepatula OGLUM05G27920 I 0.855
OLONG_022047 Oryza longistaminata OLONG_022047 I 0.855
OB05G33790 Oryza brachyantha OB05G33790 I 0.855
OB01G34490 Oryza brachyantha OB01G34490 I 0.855
LPERR05G21930 Leersia perrieri LPERR05G21930 I 0.855
mgf013423m Mimulus guttatus mgf013423m I 0.855
OBART01G25300 Oryza barthii OBART01G25300 I 0.855
OBART05G26280 Oryza barthii OBART05G26280 I 0.855
OGLUM01G29060 Oryza glumaepatula OGLUM01G29060 I 0.855
OPUNC01G25140 Oryza punctata OPUNC01G25140 I 0.855
OS01G0658900 Oryza sativa OS01G0658900 I 0.855
ORUFI05G28170 Oryza rufipogon ORUFI05G28170 I 0.855
mgf003302m Mimulus guttatus mgf003302m I 0.855
OS05G0569300 Oryza sativa OS05G0569300 I 0.855
Si001929m.g Setaria italica Si001929m.g I 0.855
ORUFI01G28110 Oryza rufipogon ORUFI01G28110 I 0.855
ORGLA05G0227800 Oryza glaberrima ORGLA05G0227800 I 0.855
ONIVA05G28290 Oryza nivara ONIVA05G28290 I 0.855
ONIVA01G28540 Oryza nivara ONIVA01G28540 I 0.855
OPUNC05G23850 Oryza punctata OPUNC05G23850 I 0.855
ORGLA01G0206000 Oryza glaberrima ORGLA01G0206000 I 0.855
LPERR01G21580 Leersia perrieri LPERR01G21580 I 0.855
OMERI03G30340 Oryza meridionalis OMERI03G30340 I 0.855
Q40645_ORYSA Oryza sativa Q40645_ORYSA D 0.855
GRMZM2G117851 Zea mays GRMZM2G117851 I 0.855
BGIOSGA001085 Oryza indica BGIOSGA001085 I 0.855
BGIOSGA020389 Oryza indica BGIOSGA020389 I 0.855
GRMZM2G011932 Zea mays GRMZM2G011932 I 0.853
Pavir.Ea02315 Panicum virgatum Pavir.Ea02315 I 0.853
GLYMA16G03190 Glycine max GLYMA16G03190 I 0.853
Sb09g028990 Sorghum bicolor Sb09g028990 I 0.853
Si022388m.g Setaria italica Si022388m.g I 0.853
BRADI2G45567 Brachypodium distachyon BRADI2G45567 I 0.853
GRMZM2G358701 Zea mays GRMZM2G358701 I 0.853
Q41109_PHAVU Phaseolus vulgaris Q41109_PHAVU D 0.853
GRMZM2G160136 Zea mays GRMZM2G160136 I 0.853
GRMZM5G848942 Zea mays GRMZM5G848942 I 0.853
GLYMA07G06620 Glycine max GLYMA07G06620 I 0.852
F775_07803 Aegilops tauschii F775_07803 I 0.851
TRIUR3_18341 Triticum urartu TRIUR3_18341 I 0.851
Traes_1BL_C5FEAF06E Triticum aestivum Traes_1BL_C5FEAF06E I 0.851
Traes_1AL_F18EC96DF Triticum aestivum Traes_1AL_F18EC96DF I 0.851
Traes_1DL_23D036A85 Triticum aestivum Traes_1DL_23D036A85 I 0.851
MLOC_6711 Hordeum vulgare MLOC_6711 I 0.851
PK09653.1 Cannabis sativa PK09653.1 D 0.850
Gorai.007G105100 Gossypium raimondii Gorai.007G105100 I 0.850
BRADI2G15940 Brachypodium distachyon BRADI2G15940 I 0.849
30174.m008698 Ricinus communis 30174.m008698 I 0.849
Ciclev10020314m.g Citrus clementina Ciclev10020314m.g I 0.845
orange1.1g014660m.g Citrus sinensis orange1.1g014660m.g I 0.845
Eucgr.E00078 Eucalyptus grandis Eucgr.E00078 I 0.844
cassava29862.valid.m1 Manihot esculenta cassava29862.valid.m1 I 0.843
cassava29861.valid.m1 Manihot esculenta cassava29861.valid.m1 I 0.843
PDK_20s1349941g001 Phoenix dactylifera PDK_20s1349941g001 I 0.841
Lus10005146.g Linum usitatissimum Lus10005146.g I 0.839
GLYMA08G34280 Glycine max GLYMA08G34280 I 0.839
Lus10003034.g Linum usitatissimum Lus10003034.g I 0.839
GSMUA_Achr6G32000_001 Musa acuminata GSMUA_Achr6G32000_001 I 0.837
TAF1_TOBAC Nicotiana tabacum TAF1_TOBAC D 0.837
gene18928-v1.0-hybrid Fragaria vesca gene18928-v1.0-hybrid I 0.836
GSMUA_Achr1G05050_001 Musa acuminata GSMUA_Achr1G05050_001 I 0.835
cassava43571.valid.m1 Manihot esculenta cassava43571.valid.m1 I 0.835
Pavir.Ca00968 Panicum virgatum Pavir.Ca00968 I 0.835
cassava43569.valid.m1 Manihot esculenta cassava43569.valid.m1 I 0.835
cassava46468.m1 Manihot esculenta cassava46468.m1 I 0.835
cassava43570.valid.m1 Manihot esculenta cassava43570.valid.m1 I 0.835
GSMUA_Achr9G05310_001 Musa acuminata GSMUA_Achr9G05310_001 I 0.834
POPTR_0002s16870 Populus trichocarpa POPTR_0002s16870 I 0.833
POPTR_0014s08990 Populus trichocarpa POPTR_0014s08990 I 0.833
GBF3 Arabidopsis lyrata fgenesh1_pm.C_scaffold_4002338 I 0.832
GBF3 Arabidopsis thaliana AT2G46270 D 0.832
Bra004550 Brassica rapa Bra004550 I 0.832
Bo4g010410 Brassica oleracea Bo4g010410 I 0.832
Carubv10023434m.g Capsella rubella Carubv10023434m.g I 0.832
Cagra.0050s0071 Capsella grandiflora Cagra.0050s0071 I 0.832
Thhalv10001533m.g Eutrema salsugineum Thhalv10001533m.g I 0.832
Ca_17787 Cicer arietinum Ca_17787 I 0.831
Pavir.J01411 Panicum virgatum Pavir.J01411 I 0.830
Pm018422 Prunus mume Pm018422 I 0.830
PRUPE_ppa006173mg Prunus persica PRUPE_ppa006173mg I 0.830
MDP0000545420 Malus domestica MDP0000545420 I 0.829
MDP0000493795 Malus domestica MDP0000493795 I 0.829
MDP0000138052 Malus domestica MDP0000138052 I 0.829
Solyc01g095460.2 Solanum lycopersicum Solyc01g095460.2 I 0.828
PGSC0003DMG400000088 Solanum tuberosum PGSC0003DMG400000088 I 0.828
Cucsa.258940 Cucumis sativus Cucsa.258940 I 0.827
Cla021500 Citrullus lanatus Cla021500 I 0.827
chr1.CM0579.280.nd Lotus japonicus chr1.CM0579.280.nd I 0.826
Ca_01293 Cicer arietinum Ca_01293 I 0.825
VIT_15s0046g01440 Vitis vinifera VIT_15s0046g01440 I 0.825
MTR_7g116890 Medicago truncatula MTR_7g116890 I 0.825
GLYMA16G15810 Glycine max GLYMA16G15810 I 0.824
Aquca_013_00209 Aquilegia coerulea Aquca_013_00209 I 0.824
SapurV1A.0013s0770 Salix purpurea SapurV1A.0013s0770 I 0.821
Gorai.008G068900 Gossypium raimondii Gorai.008G068900 I 0.820
Gorai.007G205700 Gossypium raimondii Gorai.007G205700 I 0.820
MTR_8g070820 Medicago truncatula MTR_8g070820 I 0.820
Solyc05g050220.2 Solanum lycopersicum Solyc05g050220.2 I 0.817
PGSC0003DMG400033876 Solanum tuberosum PGSC0003DMG400033876 I 0.817
Bra010572 Brassica rapa Bra010572 I 0.814
Thhalv10025733m.g Eutrema salsugineum Thhalv10025733m.g I 0.814
Aquca_037_00034 Aquilegia coerulea Aquca_037_00034 I 0.812
AMTR_s00059p00094730 Amborella trichopoda AMTR_s00059p00094730 I 0.810
VIT_02s0025g01020 Vitis vinifera VIT_02s0025g01020 I 0.809
estExt_fgenesh1_pg.C_10522 Micromonas pusilla estExt_fgenesh1_pg.C_10522 I 0.809
Cagra.5996s0003 Capsella grandiflora Cagra.5996s0003 I 0.808
GBF1 Arabidopsis lyrata scaffold_700433.1 I 0.808
GBF1 Arabidopsis thaliana AT4G36730 D 0.808
Carubv10005287m.g Capsella rubella Carubv10005287m.g I 0.808
Bostr.7867s1529 Boechera stricta Bostr.7867s1529 I 0.808
Cucsa.104970 Cucumis sativus Cucsa.104970 I 0.807
GLYMA02G06550 Glycine max GLYMA02G06550 I 0.807
Cla014415 Citrullus lanatus Cla014415 I 0.807
MTR_8g077950 Medicago truncatula MTR_8g077950 I 0.807
GLYMA16G25600 Glycine max GLYMA16G25600 I 0.807
Bra005335 Brassica rapa Bra005335 I 0.806
Pm023132 Prunus mume Pm023132 I 0.806
PRUPE_ppa006484mg Prunus persica PRUPE_ppa006484mg I 0.806
estExt_fgenesh2_pg.C_Chr_020452 Micromonas sp RCC299 estExt_fgenesh2_pg.C_Chr_020452 I 0.806
Bra011701 Brassica rapa Bra011701 I 0.805
29646.m001068 Ricinus communis 29646.m001068 I 0.805
MDP0000286846 Malus domestica MDP0000286846 I 0.805
gene13716-v1.0-hybrid Fragaria vesca gene13716-v1.0-hybrid I 0.805
Bo1g005460 Brassica oleracea Bo1g005460 I 0.805
MDP0000185553 Malus domestica MDP0000185553 I 0.805
cassava45491.m1 Manihot esculenta cassava45491.m1 I 0.805
gw1.33.173.1 Physcomitrella patens gw1.33.173.1 I 0.804
Bo3g039150 Brassica oleracea Bo3g039150 I 0.803
SELMODRAFT_135978 Selaginella moellendorffii SELMODRAFT_135978 I 0.802
MDP0000239688 Malus domestica MDP0000239688 I 0.801
Bostr.3359s0141 Boechera stricta Bostr.3359s0141 I 0.801
TCM_000436 Theobroma cacao TCM_000436 I 0.800
Pm024103 Prunus mume Pm024103 I 0.799
PRUPE_ppa020524mg Prunus persica PRUPE_ppa020524mg I 0.799
POPTR_0007s12460 Populus trichocarpa POPTR_0007s12460 I 0.798
evm.TU.supercontig_158.7 Carica papaya evm.TU.supercontig_158.7 I 0.798
POPTR_0005s12750 Populus trichocarpa POPTR_0005s12750 I 0.798
gene32022-v1.0-hybrid Fragaria vesca gene32022-v1.0-hybrid I 0.797
mgf014804m Mimulus guttatus mgf014804m I 0.796
Gorai.004G181900 Gossypium raimondii Gorai.004G181900 I 0.796
SapurV1A.0255s0070 Salix purpurea SapurV1A.0255s0070 I 0.796
mgf005151m Mimulus guttatus mgf005151m I 0.796
mgf000639m Mimulus guttatus mgf000639m I 0.796
PGSC0003DMG400012660 Solanum tuberosum PGSC0003DMG400012660 I 0.795
MDP0000231274 Malus domestica MDP0000231274 I 0.795
Solyc02g085610.2 Solanum lycopersicum Solyc02g085610.2 I 0.795
Thhalv10016740m.g Eutrema salsugineum Thhalv10016740m.g I 0.794
mgf008605m Mimulus guttatus mgf008605m I 0.794
SapurV1A.0817s0110 Salix purpurea SapurV1A.0817s0110 I 0.794
Lus10026033.g Linum usitatissimum Lus10026033.g I 0.794
Lus10014324.g Linum usitatissimum Lus10014324.g I 0.794
PDK_20s1372692g002 Phoenix dactylifera PDK_20s1372692g002 I 0.793
OPUNC02G01690 Oryza punctata OPUNC02G01690 I 0.792
Eucgr.H01256 Eucalyptus grandis Eucgr.H01256 I 0.792
Ciclev10025963m.g Citrus clementina Ciclev10025963m.g I 0.791
chr2.LjT41P23.50.nc Lotus japonicus chr2.LjT41P23.50.nc I 0.791
orange1.1g018605m.g Citrus sinensis orange1.1g018605m.g I 0.791
Eucgr.E00223 Eucalyptus grandis Eucgr.E00223 I 0.791
GBF1 Vitis vinifera VIT_04s0023g01360 I 0.791
Ca_11455 Cicer arietinum Ca_11455 I 0.791
VIT_18s0001g12120 Vitis vinifera VIT_18s0001g12120 I 0.791
Bo4g038560 Brassica oleracea Bo4g038560 I 0.791
PRUPE_ppa008024mg Prunus persica PRUPE_ppa008024mg I 0.791
Pm026586 Prunus mume Pm026586 I 0.791
VIT_02s0012g02250 Vitis vinifera VIT_02s0012g02250 I 0.791
orange1.1g036551m.g Citrus sinensis orange1.1g036551m.g I 0.790
Ciclev10001333m.g Citrus clementina Ciclev10001333m.g I 0.790
Cucsa.322110 Cucumis sativus Cucsa.322110 I 0.789
gene29546-v1.0-hybrid Fragaria vesca gene29546-v1.0-hybrid I 0.789
gw1.27.125.1 Physcomitrella patens gw1.27.125.1 I 0.789
Cla008988 Citrullus lanatus Cla008988 I 0.789
SapurV1A.0051s0250 Salix purpurea SapurV1A.0051s0250 I 0.789
MTR_5g028850 Medicago truncatula MTR_5g028850 I 0.789
LjSGA_036158.1 Lotus japonicus LjSGA_036158.1 I 0.789
PK07494.1 Cannabis sativa PK07494.1 I 0.789
GLYMA09G34170 Glycine max GLYMA09G34170 I 0.789
Cla016502 Citrullus lanatus Cla016502 I 0.789
PDK_20s1368181g001 Phoenix dactylifera PDK_20s1368181g001 I 0.789
Gorai.003G185700 Gossypium raimondii Gorai.003G185700 I 0.789
Cucsa.312800 Cucumis sativus Cucsa.312800 I 0.788
SELMODRAFT_129391 Selaginella moellendorffii SELMODRAFT_129391 I 0.788
SELMODRAFT_125734 Selaginella moellendorffii SELMODRAFT_125734 I 0.788
Bostr.23794s0549 Boechera stricta Bostr.23794s0549 I 0.787
Cagra.0993s0063 Capsella grandiflora Cagra.0993s0063 I 0.787
BZIP16 Arabidopsis thaliana AT2G35530 D 0.787
Solyc08g076100.2 Solanum lycopersicum Solyc08g076100.2 I 0.787
Traes_7DL_67C4287B2 Triticum aestivum Traes_7DL_67C4287B2 I 0.787
cassava11837.valid.m1 Manihot esculenta cassava11837.valid.m1 I 0.786
POPTR_0001s01840 Populus trichocarpa POPTR_0001s01840 I 0.786
PK23964.1 Cannabis sativa PK23964.1 D 0.786
Thhalv10007879m.g Eutrema salsugineum Thhalv10007879m.g I 0.786
cassava37043.valid.m1 Manihot esculenta cassava37043.valid.m1 I 0.786
SapurV1A.0063s0130 Salix purpurea SapurV1A.0063s0130 I 0.786
Ca_15839 Cicer arietinum Ca_15839 I 0.786
bZIP68 Arabidopsis lyrata fgenesh2_kg.1__3350__AT1G32150.1 I 0.786
POPTR_0003s09630 Populus trichocarpa POPTR_0003s09630 I 0.786
gw1.3.298.1 Physcomitrella patens gw1.3.298.1 I 0.786
30068.m002661 Ricinus communis 30068.m002661 I 0.786
BZIP68 Arabidopsis thaliana AT1G32150 D 0.786
Cagra.3359s0011 Capsella grandiflora Cagra.3359s0011 I 0.786
GLYMA01G01740 Glycine max GLYMA01G01740 I 0.785
gw2.11.550.1 Micromonas sp RCC299 gw2.11.550.1 I 0.785
CPRF3_PETCR Petroselinum crispum CPRF3_PETCR D 0.785
Aquca_002_01218 Aquilegia coerulea Aquca_002_01218 I 0.784
TCM_016606 Theobroma cacao TCM_016606 I 0.784
MLOC_79342 Hordeum vulgare MLOC_79342 I 0.784
CPRF1_PETCR Petroselinum crispum CPRF1_PETCR D 0.784
Traes_5BL_F3018E8CA Triticum aestivum Traes_5BL_F3018E8CA I 0.783
Traes_5DL_75AEE92BB Triticum aestivum Traes_5DL_75AEE92BB I 0.783
EmBP-1 Triticum aestivum Traes_5AL_9BC38814C D 0.783
Solyc02g062950.2 Solanum lycopersicum Solyc02g062950.2 I 0.783
F775_14728 Aegilops tauschii F775_14728 I 0.783
EMBP1_WHEAT Triticum aestivum EMBP1_WHEAT D 0.783