ENSONIG00000007600 (Oreochromis niloticus)
IRF

TF Information

Pfam ID Interpro ID Gene ID CIS-BP ID Sequence source Animal TF db
PF00605 (IRF) IPR001346 ENSONIG00000007600 T244273_2.00 Ensembl (2018-Dec-8) Link out

Directly determined binding motifs

Name/Motif ID Species Forward Reverse Type/Study/Study ID SR
Score		 DBD
Identity
No direct experiments

Motifs from related TFs

Name/Motif ID Species Forward Reverse Type/Study/Study ID SR
Score		 DBD
Identity
IRF6
M02254_2.00 		Meleagris gallopavo
NRNCGAWACY
RGTWTCGNYN		 PBM
Weirauch et al.(2014)
pTH9210 		0.800 0.800
IRF6
M02253_2.00 		Gallus gallus
NNMCNAAWNN
NNWTTNGKNN		 PBM
Weirauch et al.(2014)
pTH9178 		0.790 0.790
Irf6
M00176_2.00 		Mus musculus
NNNHGADASN
NSTHTCDNNN		 PBM
Badis et al.(2009)
Irf6_3803 		0.790 0.790
IRF6
M05527_2.00 		Homo sapiens
RGTWTCGNNNNNNYGAWACY
RGTWTCRNNNNNNCGAWACY		 SELEX
Yin et al.(2017)
IRF6_eDBD_HT-SELEX 		0.790 0.790
IRF6
M05529_2.00 		Homo sapiens
ACCGAAACT
AGTTTCGGT		 SELEX
Yin et al.(2017)
IRF6_FL_HT-SELEX 		0.790 0.790
IRF6
M05528_2.00 		Homo sapiens
RGTWTCGNHNNDNYGAWACY
RGTWTCRNHNNDNCGAWACY		 SELEX
Yin et al.(2017)
IRF6_eDBD_Methyl-HT-SELEX 		0.790 0.790
IRF6
M05530_2.00 		Homo sapiens
ACCGAWACY
RGTWTCGGT		 SELEX
Yin et al.(2017)
IRF6_FL_Methyl-HT-SELEX 		0.790 0.790
irf6.2
M02259_2.00 		Xenopus tropicalis
NVNHGAWACH
DGTWTCDNBN		 PBM
Weirauch et al.(2014)
pTH9302 		0.781 0.781
irf5.L
M02260_2.00 		Xenopus laevis
VNCGADACHN
NDGTHTCGNB		 PBM
Weirauch et al.(2014)
pTH9367 		0.781 0.781
Irf5
M00177_2.00 		Mus musculus
NNNHGADASH
DSTHTCDNNN		 PBM
Badis et al.(2009)
Irf5_3874 		0.762 0.762
IRF5
M02256_2.00 		Oryctolagus cuniculus
NHCGAAACYN
NRGTTTCGDN		 PBM
Weirauch et al.(2014)
pTH9272 		0.752 0.752
IRF5
M03332_2.00 		Homo sapiens
YCGAAACCGAAMCY
RGKTTCGGTTTCGR		 SELEX
Jolma et al.(2013)
IRF5_1 		0.752 0.752
IRF5
M03333_2.00 		Homo sapiens
HACCGAAACYV
BRGTTTCGGTD		 SELEX
Jolma et al.(2013)
IRF5_2 		0.752 0.752
IRF5
M05535_2.00 		Homo sapiens
NCGAAACCGAWACH
DGTWTCGGTTTCGN		 SELEX
Yin et al.(2017)
IRF5_eDBD_HT-SELEX 		0.752 0.752
IRF5
M05537_2.00 		Homo sapiens
NCGAAACCGAAACY
RGTTTCGGTTTCGN		 SELEX
Yin et al.(2017)
IRF5_FL_HT-SELEX 		0.752 0.752
IRF5
M10884_2.00 		Homo sapiens Transfac license required
 Transfac license required
 Transfac
Matys et al.(2006)
V$IRF5_Q3 		0.752 0.752
IRF5
M05536_2.00 		Homo sapiens
NYGAWACYGAWACH
DGTWTCRGTWTCRN		 SELEX
Yin et al.(2017)
IRF5_eDBD_Methyl-HT-SELEX 		0.752 0.752
IRF5
M05538_2.00 		Homo sapiens
NYGAAACCGAWACY
RGTWTCGGTTTCRN		 SELEX
Yin et al.(2017)
IRF5_FL_Methyl-HT-SELEX 		0.752 0.752
For this family, TFs with SR scores > 0.700 will likely have a similar motif

DNA Binding Domains

Protein ID Domain From To Sequence
ENSONIP00000009580 IRF 9 114

Links

Other IRF family TFs
Other Oreochromis niloticus TFs

121 Related TFs

Name Species Gene ID Motif Evidence SR
Score		 Action
AAEL006160 Aedes aegypti AAEL006160 I
AAEL006167 Aedes aegypti AAEL006167 I
AAEL007040 Aedes aegypti AAEL007040 I
AALB007752 Anopheles albimanus AALB007752 I
AALB009220 Anopheles albimanus AALB009220 I
Aan_R002212 Calypte anna Aan_R002212 I
Aan_R005895 Calypte anna Aan_R005895 I
Aan_R007315 Calypte anna Aan_R007315 I
AARA004356 Anopheles arabiensis AARA004356 I
AARA011472 Anopheles arabiensis AARA011472 I
AATE002063 Anopheles atroparvus AATE002063 I
ACEP16400 Atta cephalotes ACEP16400 I
ACEP16410 Atta cephalotes ACEP16410 I
ACHR004078 Anopheles christyi ACHR004078 I
ACHR006341 Anopheles christyi ACHR006341 I
ACOM031089 Anopheles coluzzii ACOM031089 I
ACUA016618 Anopheles culicifacies ACUA016618 I
ACYPI001442 Acyrthosiphon pisum ACYPI001442 I
ACYPI006937 Acyrthosiphon pisum ACYPI006937 I
ACYPI007541 Acyrthosiphon pisum ACYPI007541 I
ADIR005600 Anopheles dirus ADIR005600 I
AEPI003479 Anopheles epiroticus AEPI003479 I
AEPI010884 Anopheles epiroticus AEPI010884 I
AFAF000166 Anopheles farauti AFAF000166 I
Afo_R004888 Aptenodytes forsteri Afo_R004888 I
Afo_R008955 Aptenodytes forsteri Afo_R008955 I
Afo_R012718 Aptenodytes forsteri Afo_R012718 I
AFUN010423 Anopheles funestus AFUN010423 I
AGAP001094 Anopheles gambiae AGAP001094 I
AGAP002506 Anopheles gambiae AGAP002506 I
Alligator_sinensis_CCG001676.1 Alligator sinensis Alligator_sinensis_CCG001676.1 I
Alligator_sinensis_CCG010991.1 Alligator sinensis Alligator_sinensis_CCG010991.1 I
AMAM011616 Anopheles maculatus AMAM011616 I
AMAM016300 Anopheles maculatus AMAM016300 I
AMEC007551 Anopheles melas AMEC007551 I
AMEM004753 Anopheles merus AMEM004753 I
AMEM017856 Anopheles merus AMEM017856 I
AMIN002518 Anopheles minimus AMIN002518 I
AMIN005759 Anopheles minimus AMIN005759 I
Aqu1.210949 Amphimedon queenslandica Aqu1.210949 I
AQUA009970 Anopheles quadriannulatus AQUA009970 I
ASTEI07770 Anopheles stephensi ASTEI07770 I
ASTEI07771 Anopheles stephensi ASTEI07771 I
ASTEI07959 Anopheles stephensi ASTEI07959 I
BGIBMGA008906 Bombyx mori BGIBMGA008906 I
BGLB004758 Biomphalaria glabrata BGLB004758 I
Brh_R000794 Buceros rhinoceros Brh_R000794 I
Brh_R001637 Buceros rhinoceros Brh_R001637 I
CapteG142159 Capitella teleta CapteG142159 I
Cflo_03909--XP_394016.2_APIME Camponotus floridanus Cflo_03909--XP_394016.2_APIME I
Cflo_03910--XP_001121886.1_APIME Camponotus floridanus Cflo_03910--XP_001121886.1_APIME I
Cflo_03911--XP_394014.2_APIME Camponotus floridanus Cflo_03911--XP_394014.2_APIME I
Cflo_03913--XP_394013.3_APIME Camponotus floridanus Cflo_03913--XP_394013.3_APIME I
CGI_10005366 Crassostrea gigas CGI_10005366 I
CLEC011122 Cimex lectularius CLEC011122 I
Cmyd_10020455 Chelonia mydas Cmyd_10020455 I
Cmyd_10020845 Chelonia mydas Cmyd_10020845 I
Cmyd_10021516 Chelonia mydas Cmyd_10021516 I
DAPPUDRAFT_290554 Daphnia pulex DAPPUDRAFT_290554 I
DAPPUDRAFT_290557 Daphnia pulex DAPPUDRAFT_290557 I
EmuJ_000490800 Echinococcus multilocularis EmuJ_000490800 I
ENSAMEG00000005325 Ailuropoda melanoleuca ENSAMEG00000005325 I
ENSAMEG00000009223 Ailuropoda melanoleuca ENSAMEG00000009223 I
ENSAMXG00000000881 Astyanax mexicanus ENSAMXG00000000881 I
ENSAMXG00000004048 Astyanax mexicanus ENSAMXG00000004048 I
ENSAMXG00000012244 Astyanax mexicanus ENSAMXG00000012244 I
ENSAMXG00000017349 Astyanax mexicanus ENSAMXG00000017349 I
ENSAPLG00000007026 Anas platyrhynchos ENSAPLG00000007026 I
ENSAPLG00000011653 Anas platyrhynchos ENSAPLG00000011653 I
ENSAPLG00000016439 Anas platyrhynchos ENSAPLG00000016439 I
ENSBTAP00000006227-D1 Bos grunniens ENSBTAP00000006227-D1 I
ENSCAFG00000031052 Canis familiaris ENSCAFG00000031052 I
ENSCING00000002253 Ciona intestinalis ENSCING00000002253 I
ENSCPOG00000002684 Cavia porcellus ENSCPOG00000002684 I
ENSCSAG00000008703 Chlorocebus sabaeus ENSCSAG00000008703 I
ENSCSAG00000012528 Chlorocebus sabaeus ENSCSAG00000012528 I
ENSCSAVG00000004072 Ciona savignyi ENSCSAVG00000004072 I
ENSFALG00000000025 Ficedula albicollis ENSFALG00000000025 I
ENSFALG00000000470 Ficedula albicollis ENSFALG00000000470 I
ENSFALG00000001493 Ficedula albicollis ENSFALG00000001493 I
ENSGACG00000011721 Gasterosteus aculeatus ENSGACG00000011721 I
ENSGALG00000026484 Gallus gallus ENSGALG00000026484 I
ENSGALG00000026821 Gallus gallus ENSGALG00000026821 I
ENSGMOG00000010468 Gadus morhua ENSGMOG00000010468 I
ENSGMOG00000014088 Gadus morhua ENSGMOG00000014088 I
ENSLACG00000011545 Latimeria chalumnae ENSLACG00000011545 I
ENSLACG00000013776 Latimeria chalumnae ENSLACG00000013776 I
ENSLACG00000015930 Latimeria chalumnae ENSLACG00000015930 I
ENSLOCG00000002744 Lepisosteus oculatus ENSLOCG00000002744 I
ENSLOCG00000015816 Lepisosteus oculatus ENSLOCG00000015816 I
ENSMEUG00000014419 Macropus eugenii ENSMEUG00000014419 I
ENSMGAG00000014534 Meleagris gallopavo ENSMGAG00000014534 I
ENSMODG00000013941 Monodelphis domestica ENSMODG00000013941 I
ENSMPUG00000001340 Mustela putorius furo ENSMPUG00000001340 I
ENSMPUG00000012508 Mustela putorius furo ENSMPUG00000012508 I
ENSMPUG00000015804 Mustela putorius furo ENSMPUG00000015804 I
ENSNLEG00000003611 Nomascus leucogenys ENSNLEG00000003611 I
ENSNLEG00000004310 Nomascus leucogenys ENSNLEG00000004310 I
ENSNLEG00000009021 Nomascus leucogenys ENSNLEG00000009021 I
ENSOANG00000031074 Ornithorhynchus anatinus ENSOANG00000031074 I
ENSOARG00000006421 Ovis aries ENSOARG00000006421 I
ENSOARG00000013599 Ovis aries ENSOARG00000013599 I
ENSOCUG00000011739 Oryctolagus cuniculus ENSOCUG00000011739 I
ENSONIG00000001025 Oreochromis niloticus ENSONIG00000001025 I
ENSONIG00000001436 Oreochromis niloticus ENSONIG00000001436 I
ENSONIG00000004914 Oreochromis niloticus ENSONIG00000004914 I
ENSONIG00000008210 Oreochromis niloticus ENSONIG00000008210 I
ENSOPRG00000000765 Ochotona princeps ENSOPRG00000000765 I
ENSP00000300305 Macaca fascicularis ENSP00000300305 I
ENSP00000352514 Macaca fascicularis ENSP00000352514 I
ENSP00000382800 Macaca fascicularis ENSP00000382800 I
ENSP00000409227-D1 Bos grunniens ENSP00000409227-D1 I
ENSPANG00000006186 Papio anubis ENSPANG00000006186 I
ENSPANG00000007357 Papio anubis ENSPANG00000007357 I
ENSPFOG00000003997 Poecilia formosa ENSPFOG00000003997 I
ENSPFOG00000013752 Poecilia formosa ENSPFOG00000013752 I
ENSPFOG00000016115 Poecilia formosa ENSPFOG00000016115 I
ENSPPYG00000001714 Pongo abelii ENSPPYG00000001714 I
ENSPPYG00000011394 Pongo abelii ENSPPYG00000011394 I
ENSPPYG00000016664 Pongo abelii ENSPPYG00000016664 I
ENSPSIG00000012728 Pelodiscus sinensis ENSPSIG00000012728 I
ENSPSIG00000014293 Pelodiscus sinensis ENSPSIG00000014293 I
ENSPSIG00000017547 Pelodiscus sinensis ENSPSIG00000017547 I
ENSSHAG00000009950 Sarcophilus harrisii ENSSHAG00000009950 I
ENSSHAG00000012559 Sarcophilus harrisii ENSSHAG00000012559 I
ENSSHAG00000014782 Sarcophilus harrisii ENSSHAG00000014782 I
ENSSSCG00000012051 Sus scrofa ENSSSCG00000012051 I
ENSSTOG00000010163 Ictidomys tridecemlineatus ENSSTOG00000010163 I
ENSTGUG00000004719 Taeniopygia guttata ENSTGUG00000004719 I
ENSTNIG00000009855 Tetraodon nigroviridis ENSTNIG00000009855 I
ENSTNIG00000018690 Tetraodon nigroviridis ENSTNIG00000018690 I
ENSTSYG00000005097 Tarsius syrichta ENSTSYG00000005097 I
ENSXMAG00000003400 Xiphophorus maculatus ENSXMAG00000003400 I
ENSXMAG00000006715 Xiphophorus maculatus ENSXMAG00000006715 I
ENSXMAG00000018835 Xiphophorus maculatus ENSXMAG00000018835 I
frrunt Takifugu rubripes ENSTRUG00000006274 N
GA13742 Drosophila pseudoobscura FBgn0073778 I
GA22609 Drosophila pseudoobscura FBgn0244012 I
GB52719 Apis mellifera GB52719 I
GD16951 Drosophila simulans FBgn0188530 I
GD17495 Drosophila simulans FBgn0189047 I
GE17788 Drosophila yakuba FBgn0235232 I
GE17890 Drosophila yakuba FBgn0235328 I
GF15967 Drosophila ananassae FBgn0092990 I
GF15970 Drosophila ananassae FBgn0092993 I
Gfo_R000714 Geospiza fortis Gfo_R000714 I
Gfo_R011596 Geospiza fortis Gfo_R011596 I
GG18309 Drosophila erecta FBgn0110525 I
GG19691 Drosophila erecta FBgn0111893 I
GH17737 Drosophila grimshawi FBgn0125207 I
GH17740 Drosophila grimshawi FBgn0125210 I
GI14398 Drosophila mojavensis FBgn0137150 I
GI15491 Drosophila mojavensis FBgn0138240 I
GJ18742 Drosophila virilis FBgn0205894 I
GJ19245 Drosophila virilis FBgn0206390 I
GK19810 Drosophila willistoni FBgn0221808 I
GK19815 Drosophila willistoni FBgn0221813 I
GK25373 Drosophila willistoni FBgn0227332 I
GL15800 Drosophila persimilis FBgn0153404 I
GL15806 Drosophila persimilis FBgn0153410 I
GL16499 Drosophila persimilis FBgn0154103 I
GM22478 Drosophila sechellia FBgn0177348 I
GM23044 Drosophila sechellia FBgn0177912 I
GM23050 Drosophila sechellia FBgn0177918 I
Hsal_04635--XP_394013.3_APIME Harpegnathos saltator Hsal_04635--XP_394013.3_APIME I
Hsal_12319--XP_394016.2_APIME Harpegnathos saltator Hsal_12319--XP_394016.2_APIME I
Hsal_12321--XP_001121886.1_APIME Harpegnathos saltator Hsal_12321--XP_001121886.1_APIME I
ISCW004493 Ixodes scapularis ISCW004493 I
KGM_21067 Danaus plexippus KGM_21067 I
KGM_21075 Danaus plexippus KGM_21075 I
L798_10922 Zootermopsis nevadensis L798_10922 I
LLOJ002717 Lutzomyia longipalpis LLOJ002717 I
lz Drosophila melanogaster FBgn0002576 D
MCINX000302 Melitaea cinxia MCINX000302 I
MCINX005905 Melitaea cinxia MCINX005905 I
MDOA009046 Musca domestica MDOA009046 I
MDOA010744 Musca domestica MDOA010744 I
MDOA011942 Musca domestica MDOA011942 I
MDOA013149 Musca domestica MDOA013149 I
MESCA007078 Megaselia scalaris MESCA007078 I
ML03045a Mnemiopsis leidyi ML03045a I
Myotis_brandtii_RUNX2_10026084 Myotis brandtii Myotis_brandtii_RUNX2_10026084 I
NV10998 Nasonia vitripennis NV10998 I
NV11000 Nasonia vitripennis NV11000 I
NV17759 Nasonia vitripennis NV17759 I
Ogr17914 Ophisaurus gracilis Ogr17914 I
PHUM333300 Pediculus humanus PHUM333300 I
PHUM333310 Pediculus humanus PHUM333310 I
PHUM333340 Pediculus humanus PHUM333340 I
PPAI006591 Phlebotomus papatasi PPAI006591 I
PPAI010986 Phlebotomus papatasi PPAI010986 I
Pvit_01165 Pogona vitticeps Pvit_01165 I
Pvit_04762 Pogona vitticeps Pvit_04762 I
Pvit_17193 Pogona vitticeps Pvit_17193 I
RPRC005391 Rhodnius prolixus RPRC005391 I
run Drosophila melanogaster FBgn0003300 D
run Drosophila pseudoobscura FBgn0013862 I
run Drosophila virilis FBgn0013920 I
run Drosophila simulans FBgn0016352 I
run Drosophila yakuba FBgn0084618 I
run Drosophila erecta FBgn0085118 I
RUNX1 Bos taurus ENSBTAG00000004742 I
RUNX1 Canis familiaris ENSCAFG00000009596 I
RUNX1 Callithrix jacchus ENSCJAG00000002816 I
RUNX1 Dasypus novemcinctus ENSDNOG00000015041 I
Runx1 Dipodomys ordii ENSDORG00000004347 I
RUNX1 Equus caballus ENSECAG00000003462 I
RUNX1 Homo sapiens ENSG00000159216 D
RUNX1 Gasterosteus aculeatus ENSGACG00000015276 I
RUNX1 Gallus gallus ENSGALG00000016022 I
RUNX1 Gorilla gorilla ENSGGOG00000012862 I
RUNX1 Loxodonta africana ENSLAFG00000004290 I
RUNX1 Microcebus murinus ENSMICG00000011998 I
RUNX1 Macaca mulatta ENSMMUG00000001649 I
RUNX1 Monodelphis domestica ENSMODG00000021051 I
Runx1 Mus musculus ENSMUSG00000022952 D
RUNX1 Oryctolagus cuniculus ENSOCUG00000013731 I
RUNX1 Otolemur garnettii ENSOGAG00000033362 I
RUNX1 Oryzias latipes ENSORLG00000020699 I
RUNX1 Procavia capensis ENSPCAG00000016747 I
RUNX1 Pan troglodytes ENSPTRG00000013883 I
RUNX1 Pteropus vampyrus ENSPVAG00000001606 I
Runx1 Rattus norvegicus ENSRNOG00000001704 I
runx1 Takifugu rubripes ENSTRUG00000012611 I
RUNX1 Tursiops truncatus ENSTTRG00000014160 I
runx1.L Xenopus laevis XB-GENE-864949 I
RUNX2 Anolis carolinensis ENSACAG00000013003 I
RUNX2 Bos taurus ENSBTAG00000004104 I
RUNX2 Cavia porcellus ENSCPOG00000005277 I
RUNX2 Dasypus novemcinctus ENSDNOG00000034361 I
RUNX2 Equus caballus ENSECAG00000020875 I
RUNX2 Erinaceus europaeus ENSEEUG00000009243 I
RUNX2 Felis catus ENSFCAG00000022480 I
RUNX2 Homo sapiens ENSG00000124813 D
RUNX2 Gorilla gorilla ENSGGOG00000012890 I
RUNX2 Loxodonta africana ENSLAFG00000005054 I
RUNX2 Meleagris gallopavo ENSMGAG00000014267 I
RUNX2 Myotis lucifugus ENSMLUG00000000514 I
RUNX2 Macaca mulatta ENSMMUG00000019819 I
RUNX2 Monodelphis domestica ENSMODG00000018912 I
Runx2 Mus musculus ENSMUSG00000039153 D
RUNX2 Ornithorhynchus anatinus ENSOANG00000008265 I
RUNX2 Otolemur garnettii ENSOGAG00000003900 I
RUNX2 Ochotona princeps ENSOPRG00000002943 I
runx2 Oryzias latipes ENSORLG00000010169 I
RUNX2 Pan troglodytes ENSPTRG00000018228 I
Runx2 Rattus norvegicus ENSRNOG00000020193 I
RUNX2 Tupaia belangeri ENSTBEG00000000948 I
RUNX2 Taeniopygia guttata ENSTGUG00000013249 I
runx2 Takifugu rubripes ENSTRUG00000003501 I
RUNX2 Tursiops truncatus ENSTTRG00000006943 I
runx2 Xenopus tropicalis XB-GENE-482021 I
runx2.L Xenopus laevis XB-GENE-864817 I
runx2a Danio rerio ENSDARG00000040261 I
runx2b Danio rerio ENSDARG00000059233 I
runx2b Gasterosteus aculeatus ENSGACG00000012322 I
runx2b Tetraodon nigroviridis ENSTNIG00000009685 I
RUNX3 Anolis carolinensis ENSACAG00000017855 I
RUNX3 Bos taurus ENSBTAG00000019800 I
RUNX3 Callithrix jacchus ENSCJAG00000020815 I
runx3 Danio rerio ENSDARG00000052826 I
RUNX3 Dasypus novemcinctus ENSDNOG00000043493 I
runx3 Gasterosteus aculeatus ENSGACG00000007301 I
RUNX3 Loxodonta africana ENSLAFG00000008382 I
RUNX3 Myotis lucifugus ENSMLUG00000003395 I
runx3 Oryzias latipes ENSORLG00000017515 I
RUNX3 Taeniopygia guttata ENSTGUG00000001130 I
runx3 Tetraodon nigroviridis ENSTNIG00000016911 I
runx3 Takifugu rubripes ENSTRUG00000001517 I
runx3 Xenopus tropicalis XB-GENE-487784 I
RunxB Drosophila melanogaster FBgn0259162 I
Sakowv30005496m.g Saccoglossus kowalevskii Sakowv30005496m.g I
SINV13317 Solenopsis invicta SINV13317 I
SPU_006917 Strongylocentrotus purpuratus SPU_006917 I
SPU_007852 Strongylocentrotus purpuratus SPU_007852 I
TC005566 Tribolium castaneum TC005566 I
TC005796 Tribolium castaneum TC005796 I
TC006536 Tribolium castaneum TC006536 I
TC006542 Tribolium castaneum TC006542 I
tetur01g02340 Tetranychus urticae tetur01g02340 I
TriadG25179 Trichoplax adhaerens TriadG25179 I
XP_002165633.1 Hydra magnipapillata XP_002165633.1 I
XP_002939386 Xenopus tropicalis XP_002939386 I
XP_002939387 Xenopus tropicalis XP_002939387 I
XP_002939388 Xenopus tropicalis XP_002939388 I
YQE_12218 Dendroctonus ponderosae YQE_12218 I